PlantSME Plant secondary metabolism enzyme database
CAS

CAS

Triterpenoid/sterol-precursor shared module: SQS/SQE/OSC-family cyclases build the triterpenoid scaffold feeding sterol biosynthesis; overlap between terpenoid and steroid contexts is expected.

Atlas summary family CAS.

Lineage Coverage
Detected in 783 of 789 public species; click a lineage to inspect its species list.
Algae
Bryophytes
Lycophytes/Ferns
Gymnosperms
Basal angiosperms
Monocots
Eudicots
Species Distribution
Hordeum vulgare
Triticum turgidum
Aegilops tauschii
Pinus taeda
Related Modules
Steroid pathway
Pathway Position Map
Pathways are arranged by literature-curated biosynthetic stages and highlight the current enzyme family; branched pathways are grouped by stage rather than forced into one linear reaction chain.
Curated order
Steroid pathway 35857 current-family members Du et al. 2022; Zebosi et al. 2024
Sterol precursor formation
SQS/FDFT1 4389 SQE 4569 CAS 35857
Phytosterol backbone
SMT1/SMT2/SMT3 27781 SMO1/SMO2 4067 CPI1 1261 CYP51G1 190 FK 1234 HYD1 1364 DWF7 1524 DWF5 888 DWF1 1689
Brassinosteroid branch
DET2 1316 DWF4/CYP90B1 290 CPD/CYP90A1 103 ROT3/CYP90C1 193 CYP90D1 142 BR6OX1/CYP85A1 40 BR6OX2/CYP85A2 68
Family Gene List
GeneSpeciesProtein ID
arctium_lappa_Alap.ptg000011l.g3446.t1 arctium_lappa arctium_lappa_Alap.ptg000011l.g3446.t1
arctium_lappa_Alap.ptg000011l.g3447.t1 arctium_lappa arctium_lappa_Alap.ptg000011l.g3447.t1
arctium_lappa_Alap.ptg000011l.g3451.t1 arctium_lappa arctium_lappa_Alap.ptg000011l.g3451.t1
arctium_lappa_Alap.ptg000011l.g4208.t1 arctium_lappa arctium_lappa_Alap.ptg000011l.g4208.t1
arctium_lappa_Alap.ptg000012l.g1356.t1 arctium_lappa arctium_lappa_Alap.ptg000012l.g1356.t1
arctium_lappa_Alap.ptg000013l.g2350.t1 arctium_lappa arctium_lappa_Alap.ptg000013l.g2350.t1
arctium_lappa_Alap.ptg000015l.g1462.t1 arctium_lappa arctium_lappa_Alap.ptg000015l.g1462.t1
arctium_lappa_Alap.ptg000015l.g1464.t1 arctium_lappa arctium_lappa_Alap.ptg000015l.g1464.t1
arctium_lappa_Alap.ptg000015l.g1466.t1 arctium_lappa arctium_lappa_Alap.ptg000015l.g1466.t1
arctium_lappa_Alap.ptg000016l.g1073.t1 arctium_lappa arctium_lappa_Alap.ptg000016l.g1073.t1
arctium_lappa_Alap.ptg000016l.g1076.t1 arctium_lappa arctium_lappa_Alap.ptg000016l.g1076.t1
arctium_lappa_Alap.ptg000016l.g1535.t1 arctium_lappa arctium_lappa_Alap.ptg000016l.g1535.t1
arctium_lappa_Alap.ptg000016l.g1538.t1 arctium_lappa arctium_lappa_Alap.ptg000016l.g1538.t1
arctium_lappa_Alap.ptg000016l.g1539.t1 arctium_lappa arctium_lappa_Alap.ptg000016l.g1539.t1
arctium_lappa_Alap.ptg000016l.g1540.t1 arctium_lappa arctium_lappa_Alap.ptg000016l.g1540.t1
arctium_lappa_Alap.ptg000019l.g2823.t1 arctium_lappa arctium_lappa_Alap.ptg000019l.g2823.t1
arctium_lappa_Alap.ptg000020l.g1118.t1 arctium_lappa arctium_lappa_Alap.ptg000020l.g1118.t1
arctium_lappa_Alap.ptg000020l.g1562.t1 arctium_lappa arctium_lappa_Alap.ptg000020l.g1562.t1
arctium_lappa_Alap.ptg000020l.g797.t1 arctium_lappa arctium_lappa_Alap.ptg000020l.g797.t1
arctium_lappa_Alap.ptg000022l.g1006.t1 arctium_lappa arctium_lappa_Alap.ptg000022l.g1006.t1
arctium_lappa_Alap.ptg000022l.g1443.t1 arctium_lappa arctium_lappa_Alap.ptg000022l.g1443.t1
arctium_lappa_Alap.ptg000022l.g561.t1 arctium_lappa arctium_lappa_Alap.ptg000022l.g561.t1
arctium_lappa_Alap.ptg000022l.g569.t1 arctium_lappa arctium_lappa_Alap.ptg000022l.g569.t1
arctium_lappa_Alap.ptg000022l.g570.t1 arctium_lappa arctium_lappa_Alap.ptg000022l.g570.t1
arctium_lappa_Alap.ptg000022l.g571.t1 arctium_lappa arctium_lappa_Alap.ptg000022l.g571.t1
arctium_lappa_Alap.ptg000022l.g829.t1 arctium_lappa arctium_lappa_Alap.ptg000022l.g829.t1
arctium_lappa_Alap.ptg000023l.g1603.t1 arctium_lappa arctium_lappa_Alap.ptg000023l.g1603.t1
arctium_lappa_Alap.ptg000025l.g1796.t1 arctium_lappa arctium_lappa_Alap.ptg000025l.g1796.t1
arctium_lappa_Alap.ptg000025l.g2296.t1 arctium_lappa arctium_lappa_Alap.ptg000025l.g2296.t1
arctium_lappa_Alap.ptg000025l.g2298.t1 arctium_lappa arctium_lappa_Alap.ptg000025l.g2298.t1